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The hemipenis is the intromittent organ of Squamata, which is the second largest order of vertebrates with over 9,000 species distributed around the world. They differ from the intromittent organs of most other amniotes such as mammals, archosaurs and turtles that have a single genital tubercle, as squamates have the paired genitalia remaining separate. Squamate hemipenes also develop from a different cell origin, originating from the same embryonic cells that produce the li…

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